Model Internals

Each PhenDB model is trained on sets of bacterial ENOGs (orthologous groups from EggNOG 4.5), which have or have not been identified in the training genomes. Each ENOG is given a weight, with the magnitude of the weight being the importance of that ENOG for the final prediction. The sign of the weight indicates whether the presence (positive weight) or absence (negative weight) of this ENOG is indicative of the trait.

This table lists the 250 highest-ranking ENOGs of this model.

rank in model enog name enog description weight in model
1 ENOG4105DC7 peptidase M24 -0.012301
2 ENOG4105CUP Catalyzes the synthesis of alpha-ribazole-5'-phosphate from nicotinate mononucleotide (NAMN) and 5,6- dimethylbenzimidazole (DMB) (By similarity) -0.012075
3 ENOG4108UW5 maltose o-acetyltransferase 0.011727
4 ENOG4105DDQ anthranilate synthase -0.011318
5 ENOG4105EPD Oligopeptide transporter, Opt family 0.010611
6 ENOG410908R Methenyltetrahydrofolate cyclohydrolase -0.010384
7 ENOG4105M02 This protein specifically catalyzes the removal of signal peptides from prolipoproteins (By similarity) 0.010114
8 ENOG4108UHC Glycosyl transferase, wecb taga cpsf family -0.010085
9 ENOG4105C1V Peptidase, M16 0.009974
10 ENOG4107QIE Drug resistance transporter EmrB QacA -0.009882
11 ENOG4106BBN aaa ATPase 0.009680
12 ENOG4105CBD Diacylglycerol kinase -0.009609
14 ENOG4107QJP DTDP-glucose 4-6-dehydratase -0.009412
15 ENOG4107ZCT C-terminal domain of 1-Cys peroxiredoxin -0.009400
16 ENOG4108YZT Adenosylcobinamide kinase -0.009277
17 ENOG4105C0Y Responsible for the amidation of carboxylic groups at position A and C of either cobyrinic acid or hydrogenobrynic acid. NH(2) groups are provided by glutamine, and one molecule of ATP is hydrogenolyzed for each amidation (By similarity) -0.009239
18 ENOG4107QUV Domain protein -0.009187
19 ENOG4105EAG UDP-N-acetylglucosamine 2-epimerase -0.009176
20 ENOG4105DH2 Converts cobyric acid to cobinamide by the addition of aminopropanol on the F carboxylic group (By similarity) -0.009155
21 ENOG4105DYV ABC transporter 0.009131
22 ENOG4107SP8 Filamentation induced by cAMP protein fic -0.009026
23 ENOG4105C90 Catalyzes the attachment of proline to tRNA(Pro) in a two-step reaction proline is first activated by ATP to form Pro- AMP and then transferred to the acceptor end of tRNA(Pro). As ProRS can inadvertently accommodate and process non-cognate amino acids such as alanine and cysteine, to avoid such errors it has two additional distinct editing activities against alanine. One activity is designated as 'pretransfer' editing and involves the tRNA(Pro)-independent hydrolysis of activated Ala-AMP. The other activity is designated 'posttransfer' editing and involves deacylation of mischarged Ala-tRNA(Pro). The misacylated Cys- tRNA(Pro) is not edited by ProRS (By similarity) -0.008968
24 ENOG4105C5I Dehydrogenase -0.008958
25 ENOG4108UTZ Dihydroxyacetone kinase 0.008903
26 ENOG4105ESV Glycosyl transferase, family 4 -0.008898
27 ENOG4105DRP amino acid AbC transporter -0.008765
28 ENOG4105CAA catalyzes amidations at positions B, D, E, and G on adenosylcobyrinic A,C-diamide. NH(2) groups are provided by glutamine, and one molecule of ATP is hydrogenolyzed for each amidation (By similarity) -0.008740
29 ENOG4107RA1 nh(3)-dependent nad( ) synthetase -0.008663
30 ENOG4105CMT Dihydroxyacetone kinase 0.008631
31 ENOG4106538 saf domain-containing protein 0.008463
32 ENOG4108ZQX ybak prolyl-trna synthetase 0.008387
33 ENOG4108IIH CoA-transferase subunit B -0.008386
34 ENOG4108URH decarboxylase -0.008363
35 ENOG4105XBY Regulates arginine biosynthesis genes (By similarity) -0.008324
36 ENOG4105D0C transport protein 0.008295
37 ENOG4107Y6S peptidase M23 0.008272
38 ENOG4105UIB (ABC) transporter 0.008269
39 ENOG4105D7A ATP-binding protein 0.008251
40 ENOG4107YTV GtrA-like protein 0.008178
41 ENOG4105E4R Precorrin-3B C17-methyltransferase -0.008033
42 ENOG4105EN6 Membrane -0.008013
43 ENOG4105F8B mobA MobL family protein 0.007986
44 ENOG4108ZI8 NA 0.007978
45 ENOG4105CK3 Lysine 2,3-aminomutase YodO family protein -0.007858
46 ENOG4108JPW Aminotransferase -0.007835
47 ENOG4107RBQ peptidase M23 0.007831
48 ENOG4105CKD glutathione-regulated potassium-efflux system protein 0.007825
49 ENOG4108JQ9 ABC transporter 0.007803
50 ENOG4105E5K Alpha-amylase 0.007792
51 ENOG4105DBW Multi-copper polyphenol oxidoreductase laccase -0.007784
52 ENOG4105CZ1 Catalyzes the formation of N(7)-methylguanine at position 46 (m7G46) in tRNA (By similarity) 0.007756
53 ENOG4105C13 Site-specific tyrosine recombinase, which acts by catalyzing the cutting and rejoining of the recombining DNA molecules. The XerC-XerD complex is essential to convert dimers of the bacterial chromosome into monomers to permit their segregation at cell division. It also contributes to the segregational stability of plasmids (By similarity) -0.007734
54 ENOG4107QUT magnesium and cobalt transport protein CorA 0.007719
55 ENOG4105DIW FeS assembly protein SUFB -0.007714
56 ENOG4108BDP Catalyzes the transfer of endogenously produced octanoic acid from octanoyl-acyl-carrier-protein onto the lipoyl domains of lipoate-dependent enzymes. Lipoyl-ACP can also act as a substrate although octanoyl-ACP is likely to be the physiological substrate (By similarity) 0.007713
57 ENOG4105FEU K -dependent Na -Ca exchanger -0.007645
58 ENOG4107X4P Acyl-transferase 0.007637
59 ENOG4105C4B ABC transporter, permease -0.007583
60 ENOG4107QKE Glycerol-3-phosphate dehydrogenase 0.007577
61 ENOG4105CEM UPF0597 protein -0.007513
62 ENOG4108USM Transcriptional regulator, TetR family 0.007502
63 ENOG4105NI1 DNA-binding protein with PD1-like DNA-binding motif 0.007496
64 ENOG4105D98 threonine synthase 0.007477
65 ENOG4107QW6 Glycoside hydrolase family 2 TIM barrel 0.007463
66 ENOG4108ZGC gaf domain protein -0.007455
67 ENOG4105E0X reductase 0.007449
68 ENOG4107Z04 HAD-superfamily hydrolase subfamily IA 0.007331
69 ENOG4105EEM helicase -0.007285
70 ENOG4107S17 Phosphate-binding protein 0.007280
71 ENOG4105CPD ethanolamine ammonia-lyase -0.003640
71 ENOG4107FQ9 ethanolamine ammonia-lyase -0.003640
72 ENOG410811P TRANSCRIPTIONAl REGULATOR GntR family 0.007269
73 ENOG4105CVG Homoserine O-transsuccinylase -0.007265
74 ENOG4108UMA 3-octaprenyl-4-hydroxybenzoate carboxy-lyase 0.007209
75 ENOG41066HJ ATP-dependent Clp protease, proteolytic subunit 0.007200
76 ENOG4108UWE Acyltransferase -0.007194
77 ENOG4105HHQ 6-phosphogluconolactonase (EC -0.007187
78 ENOG4108RVA ribokinase 0.007185
79 ENOG4105NEH thioesterase Superfamily protein 0.007175
80 ENOG4107QU8 Transferase -0.007146
81 ENOG4105VCQ rubredoxin 0.007127
82 ENOG4105QTV Membrane 0.007121
83 ENOG4108RG0 uracil-dna glycosylase 0.007093
84 ENOG4105DN3 alpha-2-macroglobulin domain protein 0.007092
85 ENOG4105CFY cytochrome d ubiquinol oxidase, subunit ii 0.007061
86 ENOG4107R1M Catalyzes the attachment of proline to tRNA(Pro) in a two-step reaction proline is first activated by ATP to form Pro- AMP and then transferred to the acceptor end of tRNA(Pro) (By similarity) 0.007025
87 ENOG4105D7W serine acetyltransferase 0.007016
88 ENOG4105HFZ Membrane -0.006989
89 ENOG4105C54 Major Facilitator superfamily -0.006986
90 ENOG4105CG3 alpha amylase, catalytic region 0.006974
91 ENOG4108UYS had-superfamily hydrolase, subfamily ia, variant 0.006958
92 ENOG4107S5Y Glycerol-3-phosphate dehydrogenase 0.006952
93 ENOG4105ENY Ethanolamine utilization protein eutH -0.006945
94 ENOG4105CI1 PTS System 0.006927
95 ENOG4108UM1 dedA family 0.006925
96 ENOG4105DAS pts system 0.006922
97 ENOG4105F9F transposase -0.006920
98 ENOG4108UUM single-stranded DNA-binding protein 0.006875
99 ENOG4105DWV Homoserine O-trans-acetylase 0.006857
100 ENOG4108R8F zinc metallopeptidase 0.006840
101 ENOG4105CYU NDH-1 shuttles electrons from NADH, via FMN and iron- sulfur (Fe-S) centers, to quinones in the respiratory chain 0.006828
102 ENOG4108R5K PPIases accelerate the folding of proteins (By similarity) 0.006819
103 ENOG4105X1Q TOBE domain protein 0.006775
104 ENOG4108MRH transcriptional regulatory protein -0.006768
105 ENOG4107ZFB This enzyme is involved in nucleotide metabolism it produces dUMP, the immediate precursor of thymidine nucleotides and it decreases the intracellular concentration of dUTP so that uracil cannot be incorporated into DNA (By similarity) -0.006766
106 ENOG4105CUB amino acid -0.006763
107 ENOG41084AG Acylphosphatase 0.006758
108 ENOG4107QIZ NADH dehydrogenase 0.006740
109 ENOG4108UMN azlc family 0.006723
110 ENOG4105C6X Major Facilitator -0.006719
111 ENOG4105VG0 50S ribosomal protein l34 -0.006699
112 ENOG4105XEP YhcH YjgK YiaL family protein -0.006685
113 ENOG4105EE8 May catalyze the methylation of C-1 in cobalt-precorrin- 5 and the subsequent extrusion of acetic acid from the resulting intermediate to form cobalt-precorrin-6A (By similarity) -0.006675
114 ENOG4105EKT )-iron permease 0.006663
115 ENOG4108RDD Activation of anaerobic ribonucleoside-triphosphate reductase under anaerobic conditions by generation of an organic free radical, using S-adenosylmethionine and reduced flavodoxin as cosubstrates to produce 5'-deoxy-adenosine (By similarity) -0.006659
116 ENOG4105EAD ABC transporter -0.006655
117 ENOG4105C9U ATP-dependent DNA helicase RecQ -0.006651
118 ENOG4105N85 Protein of unknown function (DUF1232) 0.006625
119 ENOG4105BZ8 glutamate synthase -0.006613
120 ENOG4105EVN Prephenate dehydrogenase 0.006612
121 ENOG4105FCD Major Facilitator -0.006608
122 ENOG4108VHV Membrane 0.006597
123 ENOG4105MT2 PTS System 0.006596
124 ENOG4105CNA YD repeat protein 0.006590
125 ENOG4105FGV Chromate 0.006581
126 ENOG4108E0S reductase -0.006568
127 ENOG4108XY9 (Anaerobic) ribonucleoside-triphosphate reductase activating protein 0.006565
128 ENOG4108W0I acetyltransferase 0.006563
129 ENOG4105CK4 Destroys radicals which are normally produced within the cells and which are toxic to biological systems (By similarity) 0.006527
130 ENOG4105CV9 cyclopropane-fatty-acyl-phospholipid synthase -0.006516
131 ENOG4105D48 metallophosphoesterase 0.006507
132 ENOG4105DTW Sigma factors are initiation factors that promote the attachment of RNA polymerase to specific initiation sites and are then released (By similarity) 0.006507
133 ENOG4105CZM Major Facilitator -0.006499
134 ENOG4105MZU cell filamentation protein -0.006494
135 ENOG4107RHI Cytosine-specific methyltransferase 0.006490
136 ENOG4105CDJ oxidoreductase FAD NAD(P)-binding domain protein 0.006488
137 ENOG41076HS cation diffusion facilitator family transporter 0.006488
138 ENOG4108VY7 phosphate-starvation-inducible protein PsiE -0.006478
139 ENOG4105CK7 Sodium proline symporter -0.006465
140 ENOG4105CU5 Specifically catalyzes the decarboxylation of meso- diaminopimelate (meso-DAP) to L-lysine (By similarity) 0.006461
141 ENOG4105DNR gtp-binding protein 0.006454
142 ENOG4105K7Z Catalyzes the transfer of an acyl group from acyl- phosphate (acyl-PO(4)) to glycerol-3-phosphate (G3P) to form lysophosphatidic acid (LPA). This enzyme utilizes acyl-phosphate as fatty acyl donor, but not acyl-CoA or acyl-ACP (By similarity) 0.006438
143 ENOG4108V9Q carbonic anhydrase -0.006433
144 ENOG4105CRE cytochrome C -0.006427
145 ENOG4105KTW ethanolamine utilization protein -0.006421
146 ENOG4105CQV NDH-1 shuttles electrons from NADH, via FMN and iron- sulfur (Fe-S) centers, to quinones in the respiratory chain. The immediate electron acceptor for the enzyme in this species is believed to be ubiquinone. Couples the redox reaction to proton translocation (for every two electrons transferred, four hydrogen ions are translocated across the cytoplasmic membrane), and thus conserves the redox energy in a proton gradient (By similarity) 0.006404
147 ENOG4105WUG Toxin-antitoxin system, antitoxin component, HicB family -0.006398
148 ENOG4105CY8 phosphoribosylaminoimidazole carboxylase atpase subunit 0.006383
149 ENOG41084U9 NA 0.006364
150 ENOG4107YT9 Methyltransferase -0.006337
151 ENOG4105KT7 mgtC SapB transporter 0.006333
152 ENOG4108MDK Dehydrogenase 0.006321
153 ENOG4105W42 Veg protein 0.006299
154 ENOG4105FAV ABC, transporter 0.006293
155 ENOG4107QWA transporter -0.006283
156 ENOG4105ETU transcriptional activator (TenA -0.006281
157 ENOG41080QZ Peptidase S24-like 0.006275
158 ENOG4105U4S restriction 0.006273
159 ENOG4108IQD Involved in the biosynthesis of D-alanyl-lipoteichoic acid (LTA). Catalyzes an ATP-dependent two-step reaction where it forms a high energy D-alanyl AMP intermediate and transfers the alanyl residues from AMP to Dcp (By similarity) -0.006243
160 ENOG4108VZ1 Uncharacterised ACR, YkgG family COG1556 0.006242
161 ENOG4105K67 small basic protein -0.006229
162 ENOG4108ZEY Catalyzes the 1,3-allylic rearrangement of the homoallylic substrate isopentenyl (IPP) to its highly electrophilic allylic isomer, dimethylallyl diphosphate (DMAPP) (By similarity) -0.006222
163 ENOG4108XAN ABC transporter -0.006201
164 ENOG4105VUC UPF0237 protein -0.006201
165 ENOG4107R0S symporter 0.006195
166 ENOG4107104 thymidine kinase 0.006153
167 ENOG4107V9M epimerase dehydratase -0.006135
168 ENOG4105CWX Alpha-1,2-mannosidase 0.006133
169 ENOG4108478 ec 0.006133
170 ENOG4105D1W Converts 2,5-diamino-6-(ribosylamino)-4(3h)-pyrimidinone 5'-phosphate into 5-amino-6-(ribosylamino)-2,4(1h,3h)- pyrimidinedione 5'-phosphate (By similarity) -0.006117
171 ENOG4105CG2 Catalyzes the ATP-dependent phosphorylation of thiamine- monophosphate (TMP) to form thiamine-pyrophosphate (TPP), the active form of vitamin B1 (By similarity) 0.006109
172 ENOG4105C1B dTDP-glucose 4-6-dehydratase 0.006108
173 ENOG4105CFC Transporter -0.006101
174 ENOG41064BZ L-xylulose 5-phosphate 3-epimerase 0.006098
175 ENOG4105D5Q Arginine ornithine antiporter -0.006087
176 ENOG4105C9X ppx gppa phosphatase -0.006085
177 ENOG4108NPP Protein of unknown function (DUF1460) -0.006085
178 ENOG4105D5U fad dependent oxidoreductase 0.006083
179 ENOG4105VXR cytoplasmic protein -0.006068
180 ENOG4105DI5 UPF0365 protein 0.006061
181 ENOG4105EDK PfkB domain protein 0.006055
182 ENOG4105KMT branched-chain amino acid 0.006051
183 ENOG4105DUF Catalyzes the irreversible cleavage of the glycosidic bond in both 5'-methylthioadenosine (MTA) and S- adenosylhomocysteine (SAH AdoHcy) to adenine and the corresponding thioribose, 5'-methylthioribose and S-ribosylhomocysteine, respectively (By similarity) -0.006046
184 ENOG4105M23 ATP synthase subunit C 0.003022
184 ENOG4105X3M ATP synthase, subunit F 0.003022
185 ENOG4105K85 Involved in the cellular defense against the biological effects of O6-methylguanine (O6-MeG) in DNA. Repairs alkylated guanine in DNA by stoichiometrically transferring the alkyl group at the O-6 position to a cysteine residue in the enzyme. This is a suicide reaction the enzyme is irreversibly inactivated (By similarity) 0.006041
186 ENOG4106BHX NA 0.006038
187 ENOG4105EQ6 Hydrolyzes dipeptides containing N-terminal aspartate residues. May play a role in allowing the cell to use peptide aspartate to spare carbon otherwise required for the synthesis of the aspartate family of amino acids (By similarity) 0.006032
188 ENOG4106TCE NA 0.006024
189 ENOG4105RR1 C_GCAxxG_C_C family 0.006023
190 ENOG41085F6 50S ribosomal protein L30 0.006016
191 ENOG4105C5C n-acetylmuramoyl-l-alanine amidase 0.006011
192 ENOG4107QNU type ii secretion system protein e 0.006010
193 ENOG41084VW Part of the twin-arginine translocation (Tat) system that transports large folded proteins containing a characteristic twin-arginine motif in their signal peptide across membranes. TatA could form the protein-conducting channel of the Tat system (By similarity) 0.006005
194 ENOG4105C1K alpha-alpha-trehalose-phosphate synthase -0.005987
195 ENOG4107SEA Membrane 0.005987
196 ENOG4105FF2 Type I DHQase -0.005951
197 ENOG4105DFH Transport of potassium into the cell (By similarity) -0.005945
198 ENOG4107RRI transcriptional regulator sugar kinase 0.005944
199 ENOG4107SXS glycosyl transferase group 1 0.005940
200 ENOG4105IMH Methicillin resistance protein 0.005939
201 ENOG41082BM Transcriptional regulator, arsr family 0.005924
202 ENOG4105C5Y PTS System 0.005904
203 ENOG4105C1W two component, sigma54 specific, transcriptional regulator, Fis family 0.005904
204 ENOG410900W integral membrane protein 0.005902
205 ENOG4105D6C Part of the ABC transporter complex MacAB involved in macrolide export. Transmembrane domains (TMD) form a pore in the inner membrane and the ATP-binding domain (NBD) is responsible for energy generation (By similarity) -0.005894
206 ENOG4105NJJ -acetyltransferase 0.005892
207 ENOG4105KZP HD domain protein -0.005883
208 ENOG4105CT8 Phosphorylase 0.005881
209 ENOG4105CMZ NDH-1 shuttles electrons from NADH, via FMN and iron- sulfur (Fe-S) centers, to quinones in the respiratory chain. The immediate electron acceptor for the enzyme in this species is believed to be ubiquinone. Couples the redox reaction to proton translocation (for every two electrons transferred, four hydrogen ions are translocated across the cytoplasmic membrane), and thus conserves the redox energy in a proton gradient. This subunit may bind ubiquinone (By similarity) 0.005875
210 ENOG4108M2T Binding Domain protein -0.005872
211 ENOG4107RRN ABC transporter substrate-binding protein -0.005858
212 ENOG4105YVE ACT domain protein 0.002927
212 ENOG41069SU NA 0.002927
213 ENOG4105D1C Catalyzes the ATP- as well as the pyrophosphate- dependent phosphorylation of a specific serine residue in HPr, a phosphocarrier protein of the phosphoenolpyruvate-dependent sugar phosphotransferase system (PTS). HprK P also catalyzes the pyrophosphate-producing, inorganic phosphate-dependent dephosphorylation (phosphorolysis) of seryl-phosphorylated HPr (P- Ser-HPr). The two antagonistic activities of HprK P are regulated by several intracellular metabolites, which change their concentration in response to the absence or presence of rapidly metabolisable carbon sources (glucose, fructose, etc.) in the growth medium. Therefore, by controlling the phosphorylation state of HPr, HPrK P is a sensor enzyme that plays a major role in the regulation of carbon metabolism and sugar transport it mediates carbon catabolite repression (CCR), and regulates PTS-catalyzed carbohydrate uptake and inducer exclusion (By similarity) 0.005854
214 ENOG4105C6B Catalyzes the reversible formation of acyl-phosphate (acyl-PO(4)) from acyl- acyl-carrier-protein (acyl-ACP). This enzyme utilizes acyl-ACP as fatty acyl donor, but not acyl-CoA (By similarity) 0.005848
215 ENOG4108I66 PTS System 0.005847
216 ENOG4105CHG glycolate oxidase (iron-sulfur subunit) 0.005846
217 ENOG4105ZH6 NA 0.002921
217 ENOG4108VPR nucleoside 0.002921
218 ENOG4107SVJ Dehydrogenase -0.005838
219 ENOG4105CAG UPF0210 protein -0.005831
220 ENOG4105D60 lipoprotein releasing system transmembrane protein 0.005829
221 ENOG4108P9D Alpha beta hydrolase 0.005823
222 ENOG4105D07 Signal peptide peptidase, SppA 0.005822
223 ENOG41065VH holin, phage phi LC3 family 0.005807
224 ENOG4105FR1 rard protein -0.005801
225 ENOG4108N2V aldose 1-epimerase 0.005792
226 ENOG4107QID Threonine synthase -0.005779
227 ENOG4105C2J ABC transporter 0.005775
228 ENOG4108Z58 rRNA Methylase 0.005773
229 ENOG4105DJN thiJ pfpI -0.005763
230 ENOG4105CAJ Catalyzes the last two sequential reactions in the de novo biosynthetic pathway for UDP-N-acetylglucosamine (UDP- GlcNAc). The C-terminal domain catalyzes the transfer of acetyl group from acetyl coenzyme A to glucosamine-1-phosphate (GlcN-1-P) to produce N-acetylglucosamine-1-phosphate (GlcNAc-1-P), which is converted into UDP-GlcNAc by the transfer of uridine 5- monophosphate (from uridine 5-triphosphate), a reaction catalyzed by the N-terminal domain (By similarity) -0.005757
231 ENOG4105ZZ1 Membrane -0.005752
232 ENOG4108J0Y Diguanylate cyclase 0.005747
233 ENOG4105VD8 Resistance protein 0.005743
234 ENOG4105DSB aldose 1-epimerase 0.005720
235 ENOG4108JI7 Exporters of the RND superfamily 0.005712
236 ENOG4108ZB6 transcriptional regulator AsnC family 0.005708
237 ENOG4105D3U Ligates lysine onto the cytidine present at position 34 of the AUA codon-specific tRNA(Ile) that contains the anticodon CAU, in an ATP-dependent manner. Cytidine is converted to lysidine, thus changing the amino acid specificity of the tRNA from methionine to isoleucine (By similarity) 0.005701
238 ENOG4107T5V SNARE associated Golgi -0.005689
239 ENOG4105CIA precorrin-4 C(11)-methyltransferase -0.002836
239 ENOG41081F0 Precorrin-8x methylmutase -0.002836
240 ENOG4105DZ1 carboxypeptidase 0.005666
241 ENOG4105FF6 Modulates transcription in response to changes in cellular NADH NAD( ) redox state (By similarity) 0.005664
242 ENOG4105PAD Hydrolase -0.005656
243 ENOG4106JTU AhpC Tsa family -0.005645
244 ENOG4105FGU Nitroreductase 0.005628
245 ENOG4107R91 Catalyzes the formation of the alpha-1,6-glucosidic linkages in glycogen by scission of a 1,4-alpha-linked oligosaccharide from growing alpha-1,4-glucan chains and the subsequent attachment of the oligosaccharide to the alpha-1,6 position (By similarity) 0.005624